So, if the Karkat actually created the Pre-scratch versions of Meenah, Mituna, Porrim, Kurloz, Latula, etc. then what would be the origins of the ancestor's post scratch iterations, such as The Condense, The Psiionic, The Dolorosa, The Grand Highblood, Redglare, etc? Were they just born from the mother grub? Are they actually biologically related to the Hivebent trolls at all?

The Pre-scratch trolls and the Ancestors originated from the same wrigglers just like how the the Alpha kids and the Beta guardians are the same baby.