Well, with regards to the lips thing, some of the female trolls have different colored lips but, it's just lipstick, like how the condesce has colored lips, she kissed a panel at the end of act 6 act 5 and it came off colored, because it's lipstick. And when Vriska kissed her monitor with Nic Cage on it, and it came off blue, it's lipstick. /Trolls with colored lips are wearing lipstick./

And I mentioned before how the Megidos have red lips and the Serkets have blue lips, but I forgot to mention that while The Condence and Meenah have fuchsia lips, Feferi does not